The couple, who have been together since March 2013, walked arm-in-arm through the capital, chatting intently to each other.
Dressed in a taupe leather skirt, a blue and white striped shirt and a cream coat, actress Suki held onto Bradley's hand as he put his arm around her shoulders.
The duo's date night came after a busy day of filming for 39-year-old Bradley.
The Philadelphia-born actor was pictured in Leicester Square shooting scenes for his new film Adam Jones, which sees him take the title role of a renowned chef hoping to set up a Michelin-starred restaurant in London.
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Emma Thompson, Sienna Miller, Daniel Brühl, and Jamie Dornan make up the flick's star-studded cast as does Downton Abbey actress Lily James.
On Wednesday night Bradley and Suki dined together at London's trendiest restaurant, The Chiltern Firehouse, along with Suki's model friend Cara Delevingne.
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The couple are no strangers to the British capital, where Suki was born, and are regulars at celebrity parties, Fashion Week events and sporting competitions such as Wimbledon.